David Franks will be replaced as managing director of train operator South Central in March.
Mr Franks is moving to National Express and will be replaced by South Eastern's operations director Charles Hortin.
The reshuffle was announced last night by the Go-Ahead Group, majority partner in Govia franchises South Central and Thameslink.
Thameslink's director and general manager Mark Causebrook will become the company's managing director while Tom Smith, Govia's development director will become Govia's planning director.
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